{ "id": "fruit-development", "url": "https://coffearium.org/entries/fruit-development/", "license": "CC-BY-4.0", "title": "Fruit development", "summary": "How a fertilised coffee flower becomes a ripe cherry – the growth phases, how long each species takes, why the fruit turns red or yellow, and when the size of the bean is decided.", "domain": "botany", "kind": "Process", "stage": "plant", "topics": [ "science" ], "topicSections": {}, "lang": "en", "status": "verified", "reviewer": "Coffearium verification pass", "lastVerified": "2026-10-22T00:00:00.000Z", "updated": "2026-10-22T00:00:00.000Z", "added": "2026-10-22T00:00:00.000Z", "brief": [ "Arabica fruit grows in phases: a pinhead stage, rapid expansion, endosperm filling and ripening, which falls about 24–34 weeks after flowering.2,3", "Flowering to ripe fruit takes 6–8 months in *C. arabica* and 9–11 months in *C. canephora*, according to a 2006 review; other studies report longer spans.1,2,3", "The seed first fills with perisperm, a maternal tissue; the endosperm, the future bean, then grows into its place.1,4", "Water during rapid expansion sets the size of the space the bean can fill; shortage then gives smaller beans.2,3", "Ripe fruit turns red as chlorophyll breaks down and anthocyanins accumulate; some cultivars stay yellow.1,3" ], "quickFacts": [ { "label": "Phases (arabica)", "value": "Pinhead, rapid expansion, slow growth, endosperm filling, ripening" }, { "label": "Flower to ripe fruit, arabica", "value": "6–8 months (2006 review); 220–243 days on average (2025 review)" }, { "label": "Flower to ripe fruit, canephora", "value": "9–11 months (2006 review); 36–55 weeks (conilon clones)" }, { "label": "Short cycle", "value": "C. racemosa, 60–110 days" }, { "label": "Red colour", "value": "Anthocyanins, mainly cyanidin glycosides" }, { "label": "Critical period for bean size", "value": "Rapid expansion, about weeks 6–17" } ], "aka": [ "Coffee fruit development", "Cherry development", "Fruit growth" ], "linkTerms": [ "fruit development", "pinhead stage", "rapid expansion", "endosperm filling", "perisperm" ], "related": [ "coffee-cherry-anatomy", "flowering-and-pollination", "peaberries-and-bean-shapes", "seed-anatomy-and-storage", "coffea-arabica", "coffea-canephora", "liberica-and-excelsa", "harvesting-and-sorting", "harvest-calendars", "site-selection-and-climate", "shade-and-agroforestry", "bean-size-and-density" ], "contested": [ "Start of rapid growth in arabica: a 2007 review, after Cannell (1985), places it from about week 6 to week 17 after flowering2, and a 2025 review from 57 to 126 days3; a 2006 review cites measurements on Bourbon with rapid growth from flowering to week 7 and full size at week 171; in a 2011 Brazilian field study the fastest gain in fresh mass came only at 127–158 days, after a dry October.6", "Flower to ripe fruit in arabica: 6–8 months in a 2006 review after Cannell (1985)1; 180–250 days in a 2008 paper5; 220–243 days on average and 180–330 days overall in a 2025 review3; eight to nine months for field-grown IAPAR 59.4", "Origin of the perisperm: a 2006 review states that it develops from the nucellus of the ovule1; a 2007 review, after Cannell (1985), speaks of the expansion of the integument.2", "Why yellow-fruited arabica stays yellow: a 2025 review attributes it to leucoanthocyanin in place of anthocyanin3; a 2024 peel study, citing earlier work, attributes the colour of yellow-skinned varieties to carotenoids.8" ], "corrections": [ { "date": "2026-10-22T00:00:00.000Z", "change": "The 2025 review's flowering-to-harvest average (220–243 days) and stage limits are no longer attributed to Colombia (quick facts, body, table, chart labels, open questions).", "reason": "The review gives these values for arabica in general, citing several studies; it names Colombia only for other data." }, { "date": "2026-10-22T00:00:00.000Z", "change": "Perisperm sentence corrected: until mid-development the fruit consists mainly of pericarp and perisperm, not of perisperm alone.", "reason": "Wording of De Castro & Marraccini (2006)." } ], "sources": [ { "id": "s1", "title": "Cytology, biochemistry and molecular changes during coffee fruit development", "authors": "De Castro, R. D.; Marraccini, P.", "year": 2006, "publisher": "Brazilian Journal of Plant Physiology 18(1), 175–199", "url": "https://www.scielo.br/j/bjpp/a/hsZwcNvkPXHF97CkYgKvfKq/", "doi": "10.1590/S1677-04202006000100013", "tier": "A", "quote": "C. arabica requires 6 to 8 months to mature while C. canephora requires 9 to 11 months", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s2", "title": "Ecophysiology of coffee growth and production", "authors": "DaMatta, F. M.; Ronchi, C. P.; Maestri, M.; Barros, R. S.", "year": 2007, "publisher": "Brazilian Journal of Plant Physiology 19(4), 485–510", "url": "https://scielo.br/j/bjpp/a/xSBXnvrDDczyDbBtzspGs3v", "doi": "10.1590/S1677-04202007000400014", "tier": "A", "quote": "The expansion of the integument sets the maximum size of the bean.", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s3", "title": "Flowering and Fruiting of Coffea arabica L.: A Comprehensive Perspective from Phenology", "authors": "Unigarro, C. A.; Cayón Salinas, D. G.; León-Burgos, A. F.; Flórez-Ramos, C. P.", "year": 2025, "publisher": "Plants 14(21), 3396", "url": "https://www.mdpi.com/2223-7747/14/21/3396", "doi": "10.3390/plants14213396", "tier": "A", "quote": "this duration can vary from 180 to 330 DAF", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s4", "title": "Biochemical and genomic analysis of sucrose metabolism during coffee (Coffea arabica) fruit development", "authors": "Geromel, C.; Ferreira, L. P.; Guerreiro, S. M. C.; Cavalari, A. A.; Pot, D.; Pereira, L. F. P.; Leroy, T.; Vieira, L. G. E.; Mazzafera, P.; Marraccini, P.", "year": 2006, "publisher": "Journal of Experimental Botany 57(12), 3243–3258", "url": "https://www.alice.cnptia.embrapa.br/alice/bitstream/doc/1160617/1/Biochemical-genomic-analysis.pdf", "doi": "10.1093/jxb/erl084", "tier": "A", "quote": "fruits of C. arabica cv. IAPAR 59 completed their maturation within eight to nine months", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s5", "title": "Sucrose metabolism during fruit development in Coffea racemosa", "authors": "Geromel, C.; Ferreira, L. P.; Bottcher, A.; Pot, D.; Pereira, L. F. P.; Leroy, T.; Vieira, L. G. E.; Mazzafera, P.; Marraccini, P.", "year": 2008, "publisher": "Annals of Applied Biology 152(2), 179–187", "url": "https://www.alice.cnptia.embrapa.br/alice/bitstream/doc/880537/1/Sucrosemetabolismduring.pdf", "doi": "10.1111/j.1744-7348.2007.00199.x", "tier": "A", "quote": "was rapidly replaced by the endosperm at around 40 days after flowering (DAF)", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s6", "title": "Curvas de crescimento do fruto de cafeeiro cv. Obatã IAC 1669-20 em diferentes alinhamentos de plantio [Growth curves of coffee fruits Obatã IAC 1669-20 in different alignments of planting]", "authors": "Cunha, A. R. da; Volpe, C. A.", "year": 2011, "publisher": "Semina: Ciências Agrárias 32(1), 49–62", "url": "https://www.redalyc.org/pdf/4457/445744100005.pdf", "tier": "A", "quote": "The three alignments used showed no differences with respect the average fresh and dry mass.", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s7", "title": "Cambios físicos y químicos durante la maduración del fruto de café (Coffea arabica L. var. Colombia)", "authors": "Marín-López, S. M.; Arcila-Pulgarín, J.; Montoya-Restrepo, E. C.; Oliveros-Tascón, C. E.", "year": 2003, "publisher": "Cenicafé 54(3), 208–225", "url": "https://biblioteca.cenicafe.org/bitstream/10778/265/1/arc054(03)208-225.pdf", "tier": "A", "quote": "La calidad en taza mejor calificada se obtuvo de frutos pintones, maduros y sobremaduros (210 a 224 días después de floración), mientras que con frutos verdes (182 a 203 días) y frutos secos (231 días) fue de baja calificación.", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s8", "title": "Integrated Metabolomics and Transcriptomics Analyses Reveal the Regulatory Mechanisms of Anthocyanin and Carotenoid Accumulation in the Peel of Coffea arabica", "authors": "Wang, Z.; Xie, C.; Wu, Y.; Liu, H.; Zhang, X.; Du, H.; Li, X.; Zhang, C.", "year": 2024, "publisher": "International Journal of Molecular Sciences 25(19), 10754", "url": "https://www.mdpi.com/1422-0067/25/19/10754", "doi": "10.3390/ijms251910754", "tier": "A", "quote": "Previous studies have shown that the color of yellow-skinned varieties of Coffea arabica is attributed to carotenoids", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s9", "title": "Impacts of drought and temperature stress on coffee physiology and production: a review", "authors": "DaMatta, F. M.; Ramalho, J. D. C.", "year": 2006, "publisher": "Brazilian Journal of Plant Physiology 18(1), 55–81", "url": "https://www.scielo.br/j/bjpp/a/bDfpJwLr4xLcznSwy4b9zkf/?lang=en", "doi": "10.1590/S1677-04202006000100006", "tier": "A", "quote": "Above 23ºC, development and ripening of fruits are accelerated, often leading to loss of quality", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s10", "title": "Cultivating Coffee Quality – Terroir and Agro-Ecosystem", "authors": "Lambot, C.; Herrera, J. C.; Bertrand, B.; Sadeghian, S.; Benavides, P.; Gaitán, A.", "year": 2017, "publisher": "In: Folmer, B. (ed.), The Craft and Science of Coffee, Academic Press, ch. 2, pp. 17–49", "url": "https://doi.org/10.1016/B978-0-12-803520-7.00002-5", "doi": "10.1016/B978-0-12-803520-7.00002-5", "tier": "B", "quote": "a slower ripening process of coffee berries at higher elevations and lower air temperatures, or under shading, induces a better bean filling process", "accessed": "2026-10-22T00:00:00.000Z" }, { "id": "s11", "title": "Coffea liberica Bull ex Hiern", "authors": "Sosef, M. S. M.; Boer, E.", "year": 2000, "publisher": "In: van der Vossen, H. A. M.; Wessel, M. (eds), Plant Resources of South-East Asia No. 16: Stimulants, PROSEA Foundation, Bogor (PROSEA database record 608)", "url": "https://prosea.prota4u.org/view.aspx?id=608", "tier": "B", "quote": "red or yellow and red streaked or occasionally entirely yellow when mature", "accessed": "2026-10-22T00:00:00.000Z" } ], "markdown": "## From flower to cherry\n\nA coffee cherry starts as a fertilised flower and ends as a ripe fruit holding two seeds. How the flower opens and is pollinated is covered in [Flowering and pollination](/entries/flowering-and-pollination/). This entry follows what happens next, inside the fruit.\n\nGrowth is not steady. In arabica coffee, a 2007 review describes five stages: a pinhead stage, rapid swelling, a short pause, endosperm filling and ripening.2 A 2025 review uses the same stages with limits in days after flowering.3 Each stage builds a different part of the fruit, so a problem at one stage leaves a different mark on the harvest.\n\nThe layers of the finished fruit, from skin to seed, are described in [Coffee cherry anatomy](/entries/coffee-cherry-anatomy/). Here they matter only as they form.\n\n## The five phases\n\n**Pinhead.** For the first 6–10 weeks after flowering the young fruit hardly grows.2 It looks like a small pinhead and reaches about 10% of its final size.3\n\n**Rapid expansion.** Then the fruit swells fast, for about 10 weeks, roughly from week 6 to week 17.2 The fruit gains size and fresh mass fast in this phase.2 Inside each seed, the space is first filled by the perisperm, a tissue that comes from the mother plant.1\n\n**Slow growth.** A pause of about two weeks follows, during which the fruit reaches its final size.2\n\n**Endosperm filling.** From about week 17 to week 28, dry matter rises steadily.2 The endosperm, the tissue that becomes the green coffee bean, grows into the space the perisperm leaves behind.1 It starts as a liquid and hardens as it fills.1 The phase ends at physiological maturity: the fruit is olive green and the endosperm is solid.3\n\n**Ripening.** Ripening spreads over about 10 weeks, from week 24 to week 34 after flowering.2 The outer fruit grows again in size and mass, and turns red or yellow.2\n\n
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    \n
  1. Pinhead: the first 6–10 weeks; the fruit is about 10% of its final size and the seed space holds perisperm.2,3
    2. \n
  2. Rapid expansion: about weeks 6–17; the fruit swells to full size. In IAPAR 59 the endosperm became visible at 75 days.2,4
    3. \n
  3. Endosperm filling: about weeks 17–28; the endosperm grows into the space of the perisperm.1,2
    4. \n
  4. Physiological maturity: the fruit is olive green, the endosperm solid; only a thin layer of perisperm, the future silverskin, is left.1,3
    5. \n
  5. Ripe: about weeks 24–34; the skin turns red, or stays yellow in some cultivars.2,3
    6. \n
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An arabica fruit cut lengthwise at five stages. Hatching marks the perisperm, the shaded tissue the endosperm. Weeks after flowering after a 2007 review; the stages overlap in the sources. Schematic, not to scale.
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\n\n## What sets the size of the bean\n\nThe size of the bean is decided early, before the bean itself has formed. During rapid expansion the seed coat tissues swell and set the size of the cavity, or locule. According to a 2007 review, this expansion sets the maximum size of the bean.2 The endosperm later fills that space.1,2\n\nWater is the main limit. Water shortage during rapid expansion often reduces the growth of the fruit.2 Fruits that expand in wet weather become larger, with larger locules, than fruits that expand in hot, dry weather.2 A 2025 review links a deficit in this phase to more small beans.3\n\nSlow ripening also goes with larger beans. Longer maturation times and larger beans are usually observed at high altitude and under shade, according to a 2006 review.1 A coffee science textbook chapter explains it the same way: slower ripening allows better bean filling.10 How growers use altitude and shade is covered in [Site selection and climate](/entries/site-selection-and-climate/) and [Shade and agroforestry](/entries/shade-and-agroforestry/). Bean size as a grade is covered in [Bean size and density](/entries/bean-size-and-density/).\n\n## Green to red\n\nThe skin of the fruit stays green for most of its development.1 At the end it turns briefly yellow and then red.1 Two things happen at once: the green chlorophyll breaks down, and red pigments called anthocyanins build up.1,3 Colour is the main sign that a fruit is ripe.1\n\nSome cultivars never turn red. In natural yellow-fruited mutants of arabica the skin stays yellow when ripe.1,3 Why is not settled (see Contested). In *C. liberica*, ripe fruits are red, red-streaked yellow or occasionally fully yellow, according to a plant-resources handbook.11 How pickers use the colour stages is covered in [Harvesting and sorting](/entries/harvesting-and-sorting/).\n\n## How long each species takes\n\nThe species differ widely. A 2006 review gives 6–8 months from flowering to ripe fruit for *C. arabica* and 9–11 months for *C. canephora*.1 The same review gives more than a year for *C. liberica*, and only 10–12 weeks for *C. racemosa*, a wild species.1\n\nOther studies report longer spans for the two crop species. A 2025 review gives 220–243 days on average for arabica, and 180–330 days overall.3 Conilon clones of *C. canephora* took 36–55 weeks, depending on the clone.2 Warm weather shortens the cycle: above 23 °C, arabica fruits develop and ripen faster, often with a loss of quality.9\n\n
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Time from flowering to ripe fruit by species, as each source reports it, converted to months (our calculation, 1 month = 30.44 days). Liberica is marked at 12 months: the source says only “more than a year”.1,2,3,4,5
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\n\n\n\n## Perisperm and endosperm\n\nThe coffee seed is built twice. The perisperm develops from the nucellus of the ovule soon after fertilisation; until mid-development the fruit consists mainly of pericarp and perisperm.1 The endosperm is a triploid tissue that develops inside it.1 In *C. arabica* cv. IAPAR 59, a rapid expansion of the perisperm occurred between 60 and 90 days after flowering (DAF); the endosperm began to develop between 60 and 75 DAF and became detectable at 75 DAF.4 At 118 DAF the two tissues were present in equal proportions.4 By maturity, at 205–234 DAF, the endosperm filled the whole locule.4\n\nThe perisperm is consumed as the endosperm grows. Its inner part gives way, and in the Acaiá Cerrado measurements cited in the 2006 review the endosperm occupied the whole space by about week 19.1 Only the outer layer remains. At maturity it is a pellicle about 70 µm thick: the silverskin.1 The perisperm also stores starch for a while, which is degraded as the seed develops.4 In the endosperm, sucrose accumulated to close to 6% of dry weight.4 How the finished seed is built, and how long it stays alive, is covered in [Seed anatomy and storage](/entries/seed-anatomy-and-storage/).\n\n*C. racemosa* runs the same programme at high speed. In a field study its perisperm disappeared between 12 and 40 DAF, replaced by the endosperm, and the fruits completed their development in 68 DAF.5 The authors give 60–110 days for the species, against 180–250 days for *C. arabica*.5\n\n## Water first, dry matter later\n\nThe fruit takes up water before it stores dry matter. The 2011 field study below sampled 300 fruits of arabica cv. Obatã IAC 1669-20 every two weeks at Jaboticabal, São Paulo, Brazil.6 The authors describe slow growth from 96 to 126 DAF, rapid growth from 127 to 158 DAF and slower, linear growth from 159 to 279 DAF.6 Fresh mass grew fastest in the rapid phase, dry mass in the linear phase.6\n\n
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Fresh and dry mass of arabica fruits (cv. Obatã IAC 1669-20, Jaboticabal, Brazil, 2007/08) as a share of their highest value. Mean of six sampling positions, our calculation from Tables 2 and 3 of the source; lines only join the measured points.6
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\n\nBy our calculation from the same tables, fresh mass had reached 57% of its peak at 174 DAF, dry mass only 30%. Dry matter rose from about 17% of fresh mass during rapid growth to 31% at 279 DAF.6 The pattern matches the stage model: volume and water first, dry matter during endosperm filling.2 The timing does not. Rapid growth started about 10 weeks later than in the 2025 review, and the authors relate the slow start partly to an 83 mm water deficit in October.6 No study opened for this entry gives a full growth curve for *C. canephora* or *C. liberica*.\n\n## Bean size and water stress\n\nA 2007 review states that the expansion of the integument sets the maximum size of the bean.2 Under limiting water the ovules do not reach their potential full size.2 In a study at Ruiru, Kenya, cited in the review, potential bean size was strongly determined by rainfall 10–17 weeks after blossoming.2 The 2025 review adds that water scarcity during rapid growth can reduce the fresh mass and volume of the fruits.3\n\nLater stress does different damage. The 2025 review cites a study in which a deficit in the second part of rapid growth (about 91–119 DAF) caused black beans in 11.41% of ripe fruits.3 A continuous deficit (under 10 mm) from 57 to 80 DAF caused fruits to dry and drop, with losses of 10–40% after 80 DAF.3 These are single field results reported in a review, not general thresholds. Seeds that develop alone in a fruit, and bean shape, are covered in [Peaberries and bean shapes](/entries/peaberries-and-bean-shapes/).\n\n## Pigments of ripening\n\nThe colour change has been measured in *C. arabica* var. Colombia at 1,400 m in Chinchiná, Colombia.7 Total chlorophyll in the outer fruit tissues was 7.96–16.74 µg/g fresh weight from 182 to 203 DAF; from 210 DAF on, no chlorophyll values were reported.7 Anthocyanins rose from 1.52 µg/g at the semi-ripe stage (210 DAF) to 8.26 µg/g when ripe (217 DAF) and 47.15 µg/g when overripe (224 DAF).7 The 2006 review cites the same study with the values per gram of dry weight.1 Fruit diameter in that study grew until 217 DAF.7\n\nA 2024 study of Catimor peel found cyanidin-3-O-rutinoside, cyanidin-3-O-galactoside, cyanidin-3-O-sambubioside and cyanidin-3-O-glucoside as the main anthocyanins.8 Carotenoids, above all lutein, coloured the peel in the green and green-yellow stages; anthocyanins dominated once it turned red.8 For yellow-fruited cultivars the sources differ: the 2025 review names leucoanthocyanin in place of anthocyanin, the 2024 study carotenoids (see Contested).3,8\n\n## Reference values\n\n| Parameter | Value | Material and conditions | Source |\n|---|---|---|---|\n| Pinhead stage | First 6–10 weeks after flowering | Arabica, review after Cannell (1985) | DaMatta et al.2 |\n| Pinhead stage | Up to ≈56 DAF; ≈10% of final size | Arabica, review | Unigarro et al.3 |\n| Rapid expansion | ≈10 weeks, week 6 to week 17 | Arabica, review | DaMatta et al.2 |\n| Rapid expansion | 57–126 DAF | Arabica, review | Unigarro et al.3 |\n| Slow growth | ≈2 weeks; final size reached | Arabica, review | DaMatta et al.2 |\n| Slow growth | 126–140 DAF | Arabica, review | Unigarro et al.3 |\n| Endosperm filling | Week 17 to week 28 | Arabica, review | DaMatta et al.2 |\n| Endosperm filling | 126–196 DAF | Arabica, review | Unigarro et al.3 |\n| Ripening | Week 24 to week 34 | Arabica, review | DaMatta et al.2 |\n| Ripening | 196–224 DAF | Arabica, review | Unigarro et al.3 |\n| Endosperm first detectable | 75 DAF | Arabica cv. IAPAR 59 | Geromel et al. 20064 |\n| Perisperm and endosperm equal | 118 DAF | Arabica cv. IAPAR 59 | Geromel et al. 20064 |\n| Silverskin at maturity | ≈70 µm thick | Arabica, review | De Castro & Marraccini1 |\n| Thermal time to physiological maturity | 1,579–2,634 degree days (123–217 DAF), base 10.5 °C | Arabica cultivars, review | Unigarro et al.3 |\n| Flower to ripe fruit | 6–8 months | *C. arabica*, review after Cannell (1985) | De Castro & Marraccini1 |\n| Flower to ripe fruit | 220–243 DAF on average; 180–330 DAF overall | *C. arabica* | Unigarro et al.3 |\n| Flower to ripe fruit | Eight to nine months | *C. arabica* cv. IAPAR 59, field-grown | Geromel et al. 20064 |\n| Flower to ripe fruit | 9–11 months | *C. canephora*, review | De Castro & Marraccini1 |\n| Flower to ripe fruit | 36–55 weeks | *C. canephora* conilon clones | DaMatta et al.2 |\n| Flower to ripe fruit | More than a year | *C. liberica* | De Castro & Marraccini1 |\n| Flower to ripe fruit | 10–12 weeks; 60–110 days; 68 DAF in one field study | *C. racemosa* | De Castro & Marraccini1, Geromel et al. 20085 |\n| Anthocyanins in skin and pulp | 1.52 → 8.26 → 47.15 µg/g fresh weight at 210, 217, 224 DAF | *C. arabica* var. Colombia, 1,400 m | Marín-López et al.7 |\n| Temperature that speeds ripening | Above 23 °C | *C. arabica*, review | DaMatta & Ramalho9 |\n\nDAF = days after flowering. Weeks and days as each source gives them; the stages overlap because the sources draw their limits differently.\n\n## Open questions\n\n- **Stage timing in other regions.** The stage limits in the two reviews draw on earlier field studies they cite; the one Brazilian growth curve opened here ran about 10 weeks later.2,3,6\n- **Fruit development in canephora and liberica.** The sources opened give total durations for these species but no stage-by-stage account.1,2\n- **Yellow fruit colour.** Whether yellow-fruited arabica lacks anthocyanins because of leucoanthocyanin or because carotenoids dominate is not settled in the sources opened.3,8\n- **Maternal control of bean traits.** The 2006 review suggests that the maternal genome, through the perisperm, could be in part responsible for physical traits of the bean, though it notes that no such maternal effect had been observed in arabica or robusta breeding programmes.1\n\n## Key studies\n\n
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De Castro & Marraccini, 2006 – Cytology of coffee fruit development. A review of how the coffee fruit and seed form, from the perisperm that fills the young seed to the endosperm that replaces it and hardens. It also gathers data on colour change, storage compounds and gene expression, and compares development times across species.

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Brazilian Journal of Plant Physiology 18(1), 175–199 · doi:10.1590/S1677-04202006000100013

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DaMatta et al., 2007 – Ecophysiology of coffee growth and production. A review of how the coffee plant grows and yields under different climates. Its section on fruit growth sets out the five stages with weeks after flowering and explains why water during rapid expansion decides bean size.

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Brazilian Journal of Plant Physiology 19(4), 485–510 · doi:10.1590/S1677-04202007000400014

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Geromel et al., 2006 – Sucrose metabolism in the developing arabica fruit. Followed field-grown IAPAR 59 fruits from pinhead to maturity, separating pericarp, perisperm and endosperm. Dates when the endosperm appears and overtakes the perisperm, and tracks starch, sugars and the enzymes behind sucrose storage.

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Journal of Experimental Botany 57(12), 3243–3258 · doi:10.1093/jxb/erl084

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Geromel et al., 2008 – Fruit development in C. racemosa. Studied a wild species whose fruit matures in about two months. The perisperm is replaced within the first 40 days, which makes the species a contrast to arabica for studying seed filling.

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Annals of Applied Biology 152(2), 179–187 · doi:10.1111/j.1744-7348.2007.00199.x

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Marín-López et al., 2003 – Ripening of var. Colombia. Measured size, weight, pigments and cup quality of fruit picked at eight stages from 182 to 231 days after flowering. Chlorophyll fell and anthocyanins rose sharply in the last two weeks; the best cups came from semi-ripe to overripe fruit.

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Cenicafé 54(3), 208–225 · PDF

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